I saw a few more frames from the sequence in my morning paper. Judging from appearances - and the far-too-little time that I've been privileged to spend with these magnificent cats in Africa - I'd say it looks like this:
1) Cats are napping in their usual afternoon spot in the shade of their favorite tree. Big daddy is yawning.
2) Some human (No problem so far, they see humans all the time. They know their keepers/feeders, know where the people usually walk, know what they smell like, sound like, etc.) some human has the audacity to intrude on their space and disturb their aftenoon nap...!
3) Big daddy male lion is walking up to the edge of his customary "human border", followed by his female "backup", who does not intend to become involved (she is on alert but there is no excess muscle tension or "springing up" pose. Her ears are flattened back to indicate annoyance and the eyes are sharply focussed.)
4) Big daddy goes swat-swat with his paw while remaining seated. This means: Back off, pest. Your own house cats do this if annoyed, even if by another cat. It is simply a formal warning to behave yourself.
5) Human persists, holds out hand, instructs cat: Bite me. Cat complies, having learned from a very young age that humans are not food, but a source of food. Heck, if the lion had wanted to attack he'd have leaped right for the jugular.... Lions kill by breaking the necks of their prey.
In this case, I'm on the lions' side.